For an up-to-date list of my publications, please see my google scholar profile

International peer-reviewed publications

* denotes (at least partly) supervised students


52. Saucède T., Smith C., Olivier N., Durlet C., Gueriau P., Thoury M., Fara E., Escarguel G. & Brayard A. 2023. A new Early Triassic crinoid from Nevada questions the origin and palaeobiogeographical history of dadocrinids. Acta Palaeontologica Polonica 68, 155–166. 

51. Toussaint S.L.D., Ponstein J., Thoury M., Métivier R., Kalthoff D.C., Habermeyer B., Guilard R., Bock S., Mortensen P., Sandberg S., Gueriau P. & Amson E. 2023. Fur glowing under ultraviolet: in situ analysis of porphyrin accumulation in the skin appendages of mammals. Integrative Zoology 18, 15–26. 


50. Cupello C., Hirasawa T., Tatsumi N., Yabumoto Y., Gueriau P., Isogai S., Matsumoto R., Saruwatari T., King A., Hoshino M., Uesugi K., Okabe M. & Brito P.M. 2022Lung evolution in vertebrates and the water-to-land transition. eLife 11, e77156.

49. Celeux G., Cohen S.X., Grimaud A. & Gueriau P. 2022. Hierarchical clustering of spectral images with spatial constraints for the rapid processing of large and heterogeneous datasets. SN Computer Science 3, 194. 


48. Bertrand L., Thoury M., Gueriau P., Anheim E. & Cohen S.X. 2021. Deciphering the chemistry of cultural heritage: targeting material properties by coupling spectral imaging with image analysis. Accounts of Chemical Research 54 (13): 2823–2832.

47. Robin N., Gueriau P., Luque J., Jarvis D., Daley A.C. & Vonk R. 2021. The oldest peracarid crustacean reveals a Late Devonian freshwater colonisation by isopod relatives. Biology Letters 17 (6), 20210226

46. Alleon J., Montagnac G., Reynard B., Brule T., Thoury M., Gueriau P. 2021. Pushing Raman spectroscopy over the edge: purported signatures of organic molecules in fossil animals are instrumental artefacts. BioEssays 43 (4): 202000295.

45. Perez Peris F., Laibl L., Lustri L., Gueriau P., Antcliffe J.B., Bath Enright O. & Daley A.C. 2021. A new nektaspidid euarthropod from the Lower Ordovician of Morocco. Geological Magazine 58 (3): 509–517. 

44. Saleh F., Vidal M., Laibl L., Sansjofre P., Gueriau P., Perez Peris F., Lustri L., Lucas V., Lefebvre B., Pittet B., El Hariri K. & Daley A.C. 2021. Large trilobites in a stress-free Early Ordovician environment. Geological Magazine 58 (2): 261–270.

43. Klug C., Di Silvestro G., Hoffmann R., Schweigert G., Fuchs D., Clements T. & Gueriau P. 2021. Taphonomic patterns mimic biologic structures: Diagenetic Liesegang rings in Mesozoic coleoids and coprolites. PeerJ 9: e10703. 


42. Gueriau P., Bernard S., Farges F., Mocuta C., Dutheil D.B., Adatte T., Bomou B., Godet M., Thiaudière D., Charbonnier S. & Bertrand L. 2020. Oxidative conditions can lead to exceptional preservation through phosphatization. Geology 48 (12): 1164–1168.

41. Gueriau P., Lamsdell J.C., Wogelius R.A., Manning P.L., Egerton V.E., Bergmann U., Bertrand L. & Denayer J. 2020. A new Devonian euthycarcinoid evidences the use of different respiratory strategies during the marine-to-terrestrial transition in the myriapod lineage. Royal Society Open Science 7: 201037.

40. Gueriau P., Réguer S., Leclercq N., Cupello C., Brito P.M., Jauvion C., Morel S., Charbonnier S., Thiaudière D. & Mocuta C. 2020. Visualizing mineralization processes and fossil anatomy using synchronous synchrotron X-ray fluorescence and X-ray diffraction mapping. Journal of the Royal Society Interface 17: 20200216.

39. Cohen S.X., Webb S.M., Gueriau P., Curis E. & Bertrand L. In press. Robust framework and software implementation for fast speciation mapping. Journal of Synchrotron Radiation 27: 1049–1058 .

38. Saleh F., Pittet B., Sansjofre P., Gueriau P., Lalonde S., Perrillat J.-P., Vidal M., Lucas V., El Hariri K., Kouraiss K. & Lefebvre B. 2020. Taphonomic pathway of exceptionally preserved fossils in the Lower Ordovician of Morocco. Geobios 60: 99–115.

37. Maletz J., Mottequin B., Olive S., Gueriau P., Pernègre V., Prestianni C. & Goolaerts S. 2020. Devonian and Carboniferous dendroid graptolites from Belgium and their significance for the taxonomy of the Dendroidea. Geobios 59: 47–59.

36. Maldanis L., Hickman-Lewis K., Verezhak M., Gueriau P., Guizar-Sicairos M., Jaqueto P., Trindade R.I.F., Rossi A.L., Berenguer F., Westall F., Bertrand L. & Galante D. 2020. Nanoscale 3D quantitative imaging of 1.88 Ga Gunflint microfossils reveals novel insights into taphonomic and biogenic characters. Scientific Reports 10: 8163.

35. Jauvion C., Bernard S., Gueriau P., Mocuta C., Pont S., Benzerara K. & Charbonnier S. 2020. Exceptional preservation requires fast biodegradation: thylacocephalan specimens from La Voulte-sur-Rhône (Callovian, Jurassic, France). Palaeontology 63: 395–413 .

34. Saleh F., Antcliffe J.B., Lefebvre B., Pittet B., Laibl L., Perez Peris F., Lustri L., Gueriau P. & Daley A.C. 2020. Fossilization bias in exceptionally preserved biotas. Earth and Planetary Science Letters 529: 115873.


33. *Georgiou R., Gueriau P., Sahle C., Bernard S., Mirone A., Garrouste R., Bergmann U., Rueff J.-P. & Bertrand L. 2019. Carbon speciation in organic fossils using 2D to 3D x-ray Raman multispectral imaging. Science Advances 5: eaaw5019.

32. Klug C., Landman N.H., Fuchs D., Mapes R.H., Pohle A., Gueriau P., Réguer S. & Hoffmann R. 2019. Anatomy of the first Coleoidea and character evolution in the Carboniferous. Communications Biology 2: 280.

31. Nadhira A., Sutton M.D, Botting J.P., Muir L.A., Gueriau P., King A., Briggs D.E.G, SiveterDa.J. & Siveter De.J. 2019. Three-dimensionally preserved soft-tissues and calcareous hexactins in a Silurian sponge: implications for early sponge evolution. Royal Society Open Science 6: 190911.

30. Olive S., Pradel A., Martinez-Pérez C., Janvier P., Lamsdell J.C., Gueriau P., Rabet N., Duranleau-Gagnon P., Cardenas-Rozo A.L., Zapata Ramirez P.A. & Botella H. 2019. New insights into Late Devonian vertebrates and associated fauna from the Cuche Formation (Floresta Massif, Colombia). Journal of Vertebrate Paleontology 39: e1620247.

29. Iniesto M., Thomazo C., Fara E. & the Paris Biota team. 2019. Deciphering the exceptional preservation of the Early Triassic Paris Biota (Bear Lake County, Idaho, USA). Geobios 54: 81–93.

28. Brayard A., Gueriau P., Thoury M., Escarguel G. & the Paris Biota team. 2019. Glow in the dark: Use of synchrotron μXRF trace elemental mapping and multispectral macro-imaging on fossils from the Paris Biota (Bear Lake County, Idaho, USA). Geobios 54: 71–79.

27. Romano C., Argyriou T., Krumenacker L.J. & the Paris Biota team. 2019. Chondrichthyan teeth from the Early Triassic Paris Biota (Bear Lake County, Idaho, USA). Geobios 54: 63–70.

26. Thuy B., Escarguel G. & the Paris Biota team. 2019. A new brittle star (Ophiuroidea: Ophiodermatina) from the Early Triassic Paris Biota (Bear Lake County, Idaho, USA). Geobios 54: 55–61.

25. Saucède T., Vennin E., Fara E., Olivier N. & the Paris Biota team. 2019. A new holocrinid (Articulata) from the Paris Biota (Bear Lake County, Idaho, USA) highlights the high diversity of Early Triassic crinoids. Geobios 54: 45–53.

24. Charbonnier S., Brayard A. & the Paris Biota team. 2019. New thylacocephalans from the Early Triassic Paris Biota (Bear Lake County, Idaho, USA). Geobios 54: 37–43.

23. Brayard A., Jenks J., Bylund K.G. & the Paris Biota team. 2019. Ammonoids and nautiloids from the earliest Spathian Paris Biota and other early Spathian localities in southeastern Idaho, USA. Geobios 54: 13–36.

22. Botting J., Brayard A. & the Paris Biota team. 2019. A late-surviving Triassic protomonaxonid sponge from the Paris Biota (Bear Lake County, Idaho, USA). Geobios 54: 5–11.

21. Goutte S., Mason M.J., Antoniazzi M.M., Jared C., Merle D., Cazes L., Toledo L.F., el-Hafci H., Pallu S., Portier H., Schramm S., Gueriau P. & Thoury M. 2019. Intense bone fluorescence reveals hidden patterns in pumpkin toadlets. Scientific Reports 9: 5388.

20. Anné J., Edwards N.P., Brigidi F., Gueriau P., Harvey V.L., Geraki K., Buckley M. & Wogelius R.A. 2019. Advances in bone preservation: Identifying possible collagen preservation using sulfur speciation mapping. Palaeogeography, Palaeoclimatology, Palaeoecology 520: 181–187.

19. *Li J., Gueriau P., Bellato M., King A., Robbiola L., Thoury M., *Baillon M., Fossé C., Cohen S.X., Moulhérat C., Thomas A., Galtier P. & Bertrand L. 2019. Synchrotron-based phase mapping in corroded metals: insights from early copper-base artefacts. Analytical Chemistry 91(3): 1815–1825.

18. Lamsdell J.C., Lagebro L., Edgecombe G.D., Budd G.E. & Gueriau P. 2019. Stylonurine eurypterids from the Strud locality (Upper Devonian, Belgium): new insights into the ecology of freshwater sea scorpions. Geological Magazine 156(10): 1708–1714.


17. Gueriau P., Jauvion C. & Mocuta M. 2018. Show me your yttrium, and I will tell you who you are: implications for fossil imaging. Palaeontology 61(6): 981–990.

16. Davesne D., Gueriau P., Dutheil D.B. & Bertrand L. 2018. Exceptional preservation of a Cretaceous intestine provides a glimpse of the early ecological diversity of spiny-rayed fishes (Acanthomorpha, Teleostei). Scientific Reports 8: 8509.

15. Yang Q., Gueriau P., Charbonnier S., Ren D. & Béthoux O. 2018. A new tealliocaridid crustacean from the Late Carboniferous of North China and its biogeographic implications. Acta Palaeontologica Polonica 63(1): 111–116.

14. Gueriau P., Rabet N. & *Du Tien Hat E. 2018. The Strud crustacean fauna (Late Devonian, Belgium): updated review and palaeoecology of an early continental ecosystem. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107(2–3): 79–90.


13. Gueriau P., Rueff J.-P., Bernard S., Kaddissy J.A., Goler S., Sahle C.J., Sokaras D., Wogelius R.A., Manning P.L., Bergmann U. & Bertrand L. 2017. Non-invasive synchrotron-based X-Ray Raman scattering discriminates carbonaceous compounds in ancient and historical materials, Analytical Chemistry 89(20): 10819–10826.


12. Gueriau P., Bernard S. & Bertrand L. 2016. Advanced synchrotron characterization of paleontological specimens. Elements 12(1): 45–50.

11. Gueriau P., Rabet N., Clément G., Lagebro L., Vannier J., Briggs D.E.G., Charbonnier S., Olive S. & Béthoux O. 2016. A 365-million-year-old freshwater community reveals morphological and ecological stasis in branchiopod crustaceans. Current Biology 26(3): 383–390.

10. Denayer J., Prestianni C., Gueriau P., Olive S. & Clément G. 2016. Stratigraphy and depositional environments of the Late Famennian (Late Devonian) of Southern Belgium and characterization of the Strud locality. Geological Magazine 153(1): 112–127.


9. Gervais C., Thoury M., Réguer S., Gueriau P. & Mass J. 2015. Caution! Historic samples: Challenges in the micro spectroscopic analysis of Prussian blue and zinc white paintings. Applied Physics A 121(3): 949–955.

8. Gueriau P., Mocuta C. & Bertrand L. 2015. Cerium anomaly at microscale in fossils. Analytical Chemistry 87(17): 8827–8836.

7. Olive S., Clément G., Denayer J., Derycke C., Dupret V., Gerienne P., Gueriau P., Marion J.-M., Mottequin B. & Prestianni C. 2015. Flora and fauna from a new Famennian (Upper Devonian) locality at Becco, eastern Belgium. Geologica Belgica 18(2-4): 92–101.

6. Lagebro L., Gueriau P., Hegna T., Rabet N., Butler A. & Budd G.E. 2015. The oldest notostracan (Upper Devonian Strud locality, Belgium). Palaeontology 58(3): 497–509.


5. Gueriau P., Charbonnier S. & Clément G. 2014. Angustidontid crustaceans from the Late Devonian of Strud (Namur Province, Belgium): insights into the origin of Decapoda. Neues Jahrbuch für Geologie und Paläontologie 273(3): 327–337.

4. Gueriau P., Charbonnier S. & Clément G. 2014. First decapod crustaceans in a Late Devonian continental ecosystem. Palaeontology 57(6): 1203–1213.

3. Gueriau P., Mocuta C., Dutheil D.B., Cohen S.X., Thiaudière D., the OT1 Consortium, Charbonnier S., Clément G. & Bertrand L. 2014. Trace elemental imaging of rare earth elements discriminates tissues at microscale in flat fossils. PLoS One 9(1): e86946.


2. Garrouste R., Clément G., Nel P., Engel M.S., Grandcolas P., D’Haese, C., Lagebro L., Denayer J., Gueriau P., Lafaite P., Olive S., Prestianni C. & Nel A. 2013. Garrouste et al. reply. Nature 494: E4–E5.


1. Garrouste R., Clément G., Nel P., Engel M.S., Grandcolas P., D’Haese C., Lagebro L., Denayer J., Gueriau P., Lafaite P., Olive S., Prestianni C. & Nel A. 2012. A complete insect from the Late Devonian period. Nature 488: 82–85.

Other publications

O10. Gueriau P. 2022. Lumière sur les processus de fossilisation. In: Vigne J.-D., David B., Lecointre G., Chlous F., Clément G. & Sun J.-S. (Eds), La Terre, le vivant, les humains. Petites et grandes découvertes de l’histoire naturelle, 86–88. La Découverte. ISBN : 9782348075650.

O9. Jalil N.-E., Dutheil D. B., Gueriau P., Clément G. & Charbonnier C. 2022. Les paléoécosystèmes continentaux du Crétacé des Kem Kem. In: Charbonnier S. & De Wever P. (Eds), Paléontologie d’aujourd’hui, 56–57. EDP Sciences. ISBN : 978-2-7598-2717-6.

O8. Clément G., Denayer J., Gueriau P., Olive S., Pernègre V. & Prestianni C. 2022. L’environnement de vie des premiers tétrapodes de Belgique. In: Charbonnier S. & De Wever P. (Eds), Paléontologie d’aujourd’hui, 42–43. EDP Sciences. ISBN : 978-2-7598-2717-6.

O7. Bergmann U., Georgiou R., Gueriau P., Rueff J.-P. & Bertrand L. 2019. Nouvelles spectroscopies Raman X du carbone pour les matériaux anciens. Reflets de la Physique 63: 22–25.

O6. Gueriau P. 2018. Révéler les empreintes du paléoenvironnement. In: Gueriau P., Bertrand L., Janvier P., Deguillaume T., Thoury M., David S., Réguer S., Hordelalay C., de Parseval H., Merle D. & Scarnière O. (Eds), Lumière sur les matériaux anciens, 18–21. Réseau Canopé, Collection Éclairer. ISBN: 978-2-240-04640-6.

O5. Gueriau P. & Mondejar Fernandez J. 2016. Les premiers écosystèmes terrestres. Web article for the “Encyclopédie de l’Environnement” (Université Grenoble Alpes). URL: [also available in English]

O4. Gueriau P. The early continental aquatic arthropod fauna from the Late Devonian of Strud, Belgium: implications for terrestrialization strategies. In: Mottequin B., Denayer J., Königshof P., Prestianni C. & Olive S. (Eds), IGCP 596–SDS Symposium. Abstracts, Brussels, 20-22 septembre 2015. STRATA, Communications 16: 65.

O3. Clément G., Olive S., Gueriau P., Lagebro L., Prestianni C. & Denayer J. 2015. Assessment on the Late Devonian fauna of Strud. In : Mottequin B., Denayer J., Königshof P., Prestianni C. & Olive S. (Eds), IGCP 596-SDS Symposium. Abstracts, Brussels, 20-22 septembre 2015. STRATA, Communications 16: 35–36.

O2. Gueriau P. & Bertrand L. 2015. Deciphering exceptional preservation of fossils through trace elemental imaging. Microscopy Today 23(3): 20–25.

O1. Gueriau P. & Bertrand L. 2014. Deciphering exceptional preservation of fossils through trace elemental imaging. Microscopy and Microanalysis 20(S3): 2004–2005.